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I was just attempting to allow more air to the Crabs because I installed #100 Main jets and TOR exhaust.
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The #100 jets are sufficient compensation for the addition of the TOR mufflers and any further mods will require rejetting.
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I made some mild modifications to my air box with major performance changes. First I removed the primary chamber. The one near the battery. The results when poor mid range performance. It spit and sputtered like it was a Rich Mixture. I replace the primary chamber and drilled some small holes in the air box wear the air filter is located. The results when the same.
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By removing the airbox and/or drilling the filter box you've driven your mixture excessively lean. The performance symptoms are the same for either a lean or rich condition -- loss of power and misfiring in extreme cases.
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"How does changing the air flow affect the Vacuum Piston Assembly?"
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The carb slide reacts to the
velocity of the air flow through the carb throat and the velocity through the carb throat is determined by the pressure drops in the system.
Every physical element in the airflow path provides a restriction to airflow and introduces a pressure drop across that element. Each pressure drop is also a velocity drop.
Opening up the filter box and/or the inlet system reduces the inlet pressure/velocity drop(s), increases the pressure differential across the carb and the flow across the carb slide increases.
That causes the slide to open faster and the mixture goes lean because the main jet can't supply sufficient fuel for the increased airflow.
I'd suggest you either apply some duct tape over the holes you've drilled or prepare for a lot of carb removals and jet changes.
Jim